Nero is one of the main protagonists of the Devil May Cry series. He first appeared as a playable character in Devil May Cry 4, and later in Devil May Cry 5. He was raised in Fortuna and served as a Holy Knight in the Order of the Sword, a religious group that worships Sparda and fights to protect the world from demons. It is later revealed that he is the son of Vergil, nephew of Legendary Devil Hunter Dante, and grandson of Legendary Dark Knight Sparda. In Devil May Cry 4, however, he is forced to fight against the Order after encountering Dante and uncovering a conspiracy to conquer the world using The Savior. In Devil May Cry 5, Nero is a more seasoned demon hunter who has since changed his operation to being a mobile Devil May Cry branch. He finds himself fending off a demonic invasion as he chases a mysterious figure (later revealed to be his own biological father Vergil at the end) that severed the Devil Bringer.